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Francis Ruddle, Who Led Transgenic Research, Dies at 83

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Francis Ruddle, a Yale University geneticist who plumbed the DNA of mice and men, and from whose laboratory sprang the first transgenic mammal — a mouse with added foreign DNA — died on March 10 in New Haven. He was 83. Enlarge This Image Ken Laffal Francis Ruddle, in 1981, a year after his lab said it had implanted foreign DNA into mice, altering their genetic makeup. Connect With Us on Social Media @nytimesscience on Twitter. Science Reporters and Editors on Twitter Like the science desk on Facebook. The cause was complications of a heart condition, said his wife, Nancy Ruddle. Dr. Ruddle, who at his death was the Sterling professor emeritus of molecular, cellular and...

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